Seed dispersal and predation by scatter-hoarding rodents play an important role in seedling regeneration, community structure and composition, and species coexistence (Vander Wall 2010;Zhang et al.2016).Rodents generally exhibit seed species preference during their scatter-hoarding behavioral processes (Xiao et al.2006;Gong et al.2015;Zhang et al.2016), and seed traits are a key factor in explaining rodent foraging preference (Wang and Chen 2009;Vander Wall2010;Zhang et al.2016).