Breeding crops with enhanced nutrients is one of the major goals of plant breeding.Notably,plants intensively produce various specialized plant metabolites(e.g.,phenolic compounds and ca-rotenoids)in response to diverse external stimuli and changing environmental conditions,which have been reported as health-promoting compounds with various health benefits(Dong and Lin,2021).On the other hand,the varied growth environment and global climate change necessitates crops to have the qualities of good environmental adaptability(Eshed and Lippman,2019).