In the first paragraph of'On the origin of species', Darwin [1] refers to speciation as that'mystery of mysteries'.Although the means by which new species arise is no longer mysterious, some puzzles remain.One such puzzle, which is addressed in a recent paper from Chung-I Wu's group on mangrove speciation [2],concerns the popular allopatric speciation model.In this model,which is widely viewed as the most common form of speciation, geographically isolated populations diverge to the point at which successful reproduction between them is no longer possible.