The macronutrient phosphorus(P)is essential for plant growth and of vital importance to crop yield.It is taken up from the soil in the form of inorganic orthophosphate[PO43-,HPO42-,H2PO4-(Pi)]by plant roots.As Pi readily forms insoluble com-plexes and precipitates with organic matter and mineral cations,Pi bioavailability in the soil is usually low and often limits crop growth and final yield.To sustain crop yields,consumption of inorganic P fertilizer derived from mining of non-renewable phos-phate rock is heavily relied upon for agriculture,which causes is-sues of P scarcity and environmental problems(Cong et al.,2020).Therefore,improving crop yields with minimal P fertilizer input is a major challenge for plant scientists.