Global temperature has been increasing at unprecedented rates during the Anthropocene,impacting both natural and human sys-tems [1].Alpine biomes,among the most sensitive natural ecosys-tems to climate warming,show rapid shifts of species distribution ranges and modulations of species interactions [1].The Himalayas(also known as the "water tower" of Asia and a global biodiversity hot spot) are highly sensitive and vulnerable to global warming as this is one of the fastest-warming regions in the world.Such rapid warming is expected to trigger upward shifts of alpine vegetation,because cold temperature limitations on growth and recruitment are being alleviated [2].However,increasing drought stress may dampen or even reverse this positive response of alpine ecosys-tems to warming climate [3A].In addition,interactions between alpine plants co-determine the structure and function of subnival vegetation,and thereby stabilize their distribution range [5].