NF-YB1-regulated expression of sucrose transporters in aleurone facilitates sugar loading to rice endosperm
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摘要:
Dear Editor,
Cereal endosperms, which consist of an outer aleurone layer and an inner starchy endosperm, provide over 70% staple food for human [www.fao.org].Rice endosperm development occurs in the caryopsis of a grain.During the grain filling stage, sucrose produced in photosyntheticleaves is transported to grains via the phloem [1, 2].It has been proposed that sucrose, upon arrival in the caryopsis through the phloem located at the dorsal vascular bundle, is partially hydrolyzed by cell wall invertases (CINs) into glucose and fructose [3, 4].Since endosperm is symplastically isolated from maternal tissues, it is assumed that these sugars are delivered to the endosperm actively through an apoplastic pathway by sugar transporters located in aleurone [5,6].However, the exact role of aleurone in regulating sugar transport remains unknown.