Heterosis is a phenomenon where hybrids have superior perfor-mance over their parents.As an efficient way to increase crop yield,heterosis is of great importance on agriculture production.Therefore,hybrid breeding has been widely used in maize and rice for several decades,where the elite hybrid is developed by crossing two inbred lines(Liu et al.,2020).However,in many outcrossing plants,it is not easy to create genetic diverse inbred lines of both high genomic homozygosity and vigor due to self-incompatibility and inbreeding depression(Chen et al.,2021).